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ABSTRACT SUPPLIED BY ORIGINATOR: During the Mid-Pleistocene Transition (MPT; 1,200-800 kya), Earth&apos;s orbitally paced ice age cycles intensified, lengthened from ~40,000 (~40 ky) to ~100 ky, and became distinctly asymmetrical. Testing hypotheses that implicate changing atmospheric CO2 levels as a driver of the MPT has proven difficult with available observations. Here, we use orbitally resolved, boron isotope CO2 data to show that the glacial to interglacial CO2 difference increased from ~43 to ~75 uatm across the MPT, mainly because of lower glacial CO2 levels. Through carbon cycle modeling, we attribute this decline primarily to the initiation of substantive dust-borne iron fertilization of the Southern Ocean during peak glacial stages. We also observe a twofold steepening of the relationship between sea level and CO2-related climate forcing that is suggestive of a change in the dynamics that govern ice sheet stability, such as that expected from the removal of subglacial regolith or interhemispheric ice sheet phase-locking. We argue that neither ice sheet dynamics nor CO2 change in isolation can explain the MPT. Instead, we infer that the MPT was initiated by a change in ice sheet dynamics and that longer and deeper post-MPT ice ages were sustained by carbon cycle feedbacks related to dust fertilization of the Southern Ocean as a consequence of larger ice sheets.</gco:CharacterString>
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           <gmd:city>
            <gco:CharacterString>Asheville</gco:CharacterString>
           </gmd:city>
           <gmd:administrativeArea>
            <gco:CharacterString>NC</gco:CharacterString>
           </gmd:administrativeArea>
           <gmd:postalCode>
            <gco:CharacterString>28801-5001</gco:CharacterString>
           </gmd:postalCode>
           <gmd:country>
            <gco:CharacterString>USA</gco:CharacterString>
           </gmd:country>
           <gmd:electronicMailAddress>
            <gco:CharacterString>ncei.orders@noaa.gov</gco:CharacterString>
           </gmd:electronicMailAddress>
          </gmd:CI_Address>
         </gmd:address>
         <gmd:onlineResource>
          <gmd:CI_OnlineResource>
           <gmd:linkage>
            <gmd:URL>https://www.ncei.noaa.gov</gmd:URL>
           </gmd:linkage>
           <gmd:protocol>
            <gco:CharacterString>HTTPS</gco:CharacterString>
           </gmd:protocol>
           <gmd:applicationProfile>
            <gco:CharacterString>Web Browser</gco:CharacterString>
           </gmd:applicationProfile>
           <gmd:name>
            <gco:CharacterString>NOAA National Centers for Environmental Information (NCEI)</gco:CharacterString>
           </gmd:name>
           <gmd:description>
            <gco:CharacterString>NCEI home page with information, data access and contact information.</gco:CharacterString>
           </gmd:description>
           <gmd:function>
            <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="information">information</gmd:CI_OnLineFunctionCode>
           </gmd:function>
          </gmd:CI_OnlineResource>
         </gmd:onlineResource>
         <gmd:hoursOfService>
          <gco:CharacterString>8:00 - 6:00 Eastern</gco:CharacterString>
         </gmd:hoursOfService>
        </gmd:CI_Contact>
       </gmd:contactInfo>
       <gmd:role>
        <gmd:CI_RoleCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_RoleCode" codeListValue="distributor">distributor</gmd:CI_RoleCode>
       </gmd:role>
      </gmd:CI_ResponsibleParty>
     </gmd:distributorContact>
     <gmd:distributionOrderProcess>
      <gmd:MD_StandardOrderProcess>
       <gmd:fees>
        <gco:CharacterString>In most cases, electronic downloads of the data are free, however fees may apply for data certifications, copies of analog materials, and data distribution on physical media.</gco:CharacterString>
       </gmd:fees>
      </gmd:MD_StandardOrderProcess>
     </gmd:distributionOrderProcess>
     <gmd:distributorFormat>
      <gmd:MD_Format>
       <gmd:name>
        <gco:CharacterString>Native Format(s)</gco:CharacterString>
       </gmd:name>
       <gmd:version gco:nilReason="missing"></gmd:version>
      </gmd:MD_Format>
     </gmd:distributorFormat>
     <gmd:distributorTransferOptions>
      <gmd:MD_DigitalTransferOptions>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://data.nodc.noaa.gov/cgi-bin/iso?id=noaa-forcing-23550</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTPS</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Landing Page</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI landing page for this dataset</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Dataset landing page with general information and links to access the data.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="information">information</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www.ncdc.noaa.gov/paleo/study/23550</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTPS</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Web Browser</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Study Page</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Study page with dataset access information.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="search">search</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www.ncdc.noaa.gov/data-access/paleoclimatology-data/datasets/climate-forcing</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTPS</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Information</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Forcing Page</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>General information for this data type.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="information">information</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/climate_forcing/trace_gases/Paleo-pCO2/chalk2017odp999boron.xlsx</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/climate_forcing/trace_gases/Paleo-pCO2/chalk2017odp999boron.txt</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
      </gmd:MD_DigitalTransferOptions>
     </gmd:distributorTransferOptions>
    </gmd:MD_Distributor>
   </gmd:distributor>
  </gmd:MD_Distribution>
 </gmd:distributionInfo>
 <gmd:metadataMaintenance>
  <gmd:MD_MaintenanceInformation>
   <gmd:maintenanceAndUpdateFrequency>
    <gmd:MD_MaintenanceFrequencyCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#MD_MaintenanceFrequencyCode" codeListValue="asNeeded">asNeeded</gmd:MD_MaintenanceFrequencyCode>
   </gmd:maintenanceAndUpdateFrequency>
   <gmd:maintenanceNote>
    <gco:CharacterString>This metadata record was translated from DIF using dif-9x-to-iso-19115-2.xsl Version: 2018-10-25</gco:CharacterString>
   </gmd:maintenanceNote>
  </gmd:MD_MaintenanceInformation>
 </gmd:metadataMaintenance>
</gmi:MI_Metadata>