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ABSTRACT SUPPLIED BY ORIGINATOR: The transport of moisture in the tropics is a critical process for the global energy budget and on geologic timescales, has markedly influenced continental landscapes, migratory pathways, and biological evolution. Here we present a continuous, first-of-its-kind 1.3-My record of continental hydroclimate and lake-level variability derived from drill core data from Lake Malawi, East Africa (9-15S). Over the Quaternary, we observe dramatic shifts in effective moisture, resulting in large-scale changes in one of the world&apos;s largest lakes and most diverse freshwater ecosystems. Results show evidence for 24 lake level drops of more than 200 m during the Late Quaternary, including 15 lowstands when water levels were more than 400 m lower than modern. A dramatic shift is observed at the Mid-Pleistocene Transition (MPT), consistent with far-field climate forcing, which separates vastly different hydroclimate regimes before and after ~800,000 years ago. Before 800 ka, lake levels were lower, indicating a climate drier than today, and water levels changed frequently. Following the MPT high-amplitude lake level variations dominate the record. From 800 to 100 ka, a deep, often overfilled lake occupied the basin, indicating a wetter climate, but these highstands were interrupted by prolonged intervals of extreme drought. Periods of high lake level are observed during times of high eccentricity. The extreme hydroclimate variability exerted a profound influence on the Lake Malawi endemic cichlid fish species flock; the geographically extensive habitat reconfiguration provided novel ecological opportunities, enabling new populations to differentiate rapidly to distinct species.</gco:CharacterString>
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          <gco:CharacterString>Landing Page</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI landing page for this dataset</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Dataset landing page with general information and links to access the data.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="information">information</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www.ncdc.noaa.gov/paleo/study/19424</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTPS</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Web Browser</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Study Page</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Study page with dataset access information.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="search">search</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www.ncdc.noaa.gov/data-access/paleoclimatology-data/datasets/lake</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTPS</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Information</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Lake Page</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>General information for this data type.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="information">information</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/paleolimnology/eastafrica/malawi2015pca.txt</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/paleolimnology/eastafrica/malawi2015carb.txt</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/paleolimnology/eastafrica/malawi2015color.txt</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/paleolimnology/eastafrica/malawi2015density.txt</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/paleolimnology/eastafrica/malawi2015ca.txt</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
       <gmd:onLine>
        <gmd:CI_OnlineResource>
         <gmd:linkage>
          <gmd:URL>https://www1.ncdc.noaa.gov/pub/data/paleo/paleolimnology/eastafrica/malawi2015.xlsx</gmd:URL>
         </gmd:linkage>
         <gmd:protocol>
          <gco:CharacterString>HTTP</gco:CharacterString>
         </gmd:protocol>
         <gmd:applicationProfile>
          <gco:CharacterString>Download</gco:CharacterString>
         </gmd:applicationProfile>
         <gmd:name>
          <gco:CharacterString>NCEI Direct Download</gco:CharacterString>
         </gmd:name>
         <gmd:description>
          <gco:CharacterString>Direct download links for dataset files.</gco:CharacterString>
         </gmd:description>
         <gmd:function>
          <gmd:CI_OnLineFunctionCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#CI_OnLineFunctionCode" codeListValue="download">download</gmd:CI_OnLineFunctionCode>
         </gmd:function>
        </gmd:CI_OnlineResource>
       </gmd:onLine>
      </gmd:MD_DigitalTransferOptions>
     </gmd:distributorTransferOptions>
    </gmd:MD_Distributor>
   </gmd:distributor>
  </gmd:MD_Distribution>
 </gmd:distributionInfo>
 <gmd:metadataMaintenance>
  <gmd:MD_MaintenanceInformation>
   <gmd:maintenanceAndUpdateFrequency>
    <gmd:MD_MaintenanceFrequencyCode codeList="https://data.noaa.gov/resources/iso19139/schema/resources/Codelist/gmxCodelists.xml#MD_MaintenanceFrequencyCode" codeListValue="asNeeded">asNeeded</gmd:MD_MaintenanceFrequencyCode>
   </gmd:maintenanceAndUpdateFrequency>
   <gmd:maintenanceNote>
    <gco:CharacterString>This metadata record was translated from DIF using dif-9x-to-iso-19115-2.xsl Version: 2018-10-25</gco:CharacterString>
   </gmd:maintenanceNote>
  </gmd:MD_MaintenanceInformation>
 </gmd:metadataMaintenance>
</gmi:MI_Metadata>